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Plant

Cactus

Cactaceae (family)

Also known as: Cactaceae

A family of around 1,750 species of succulent flowering plants — almost all native to the Americas, with the single exception of *Rhipsalis baccifera*, which also occurs in Africa, Madagascar, and Sri Lanka (likely via natural seed dispersal across the Atlantic by birds in the geologically recent past). Cacti are the New World's convergent evolutionary answer to the Old World's [[agave]], [[aloe-vera]], and *Euphorbia* species — different families that have separately evolved the same suite of adaptations to extreme aridity. The visual signature of the American Southwest, the Andean altiplano, the Mexican deserts, and a substantial chunk of modern horticultural houseplant culture.

Cactus
Photo via Wikimedia Commons — see source for license.

Scientific

Cactaceae contains ~1,750 species across approximately 130 genera. Notable genera include:

  • Carnegiea — the [[saguaro|saguaro]]; covered separately at [[saguaro]]
  • Opuntia — [[prickly-pear|prickly pear]]; widely naturalized globally; the nopal of Mexican cuisine
  • Mammillaria — small spherical “pincushion” cacti; among the most-cultivated houseplants
  • Echinocactus — barrel cacti
  • Cereus, Pachycereus — large columnar cacti
  • Astrophytum — geometric “star” cacti
  • Rhipsalis — the unusual epiphytic forest cacti; the genus including the only naturally Old-World-distributed cactus species

Cactaceae’s defining features:

  • Areoles — small cushion-like structures from which spines, flowers, and new growth emerge; unique to Cactaceae
  • Succulent stems — water storage in the stem rather than the leaves
  • CAM photosynthesis — Crassulacean Acid Metabolism, where stomata open at night (to avoid daytime water loss) and CO2 is fixed in malic acid for daytime use

Spines are highly modified leaves — over evolutionary time, leaves were reduced (and then eliminated in most species) in favor of stem-based water storage and CAM photosynthesis. The spines that remain serve dual roles: defense against herbivores and reduction of solar heat-load on the stem through shading.

Convergent evolution

Cacti’s adaptations are evolutionary parallels — not relations — of similar adaptations in Old World succulents:

  • [[Aloe-vera]] and other aloes (Asphodelaceae) — Africa
  • [[Agave]] (Asparagaceae) — Americas, but in a different family from cacti
  • Euphorbia succulents (Euphorbiaceae) — Africa, especially southern Africa

The cactus / euphorbia confusion is famous in horticulture — many Euphorbia succulents look indistinguishable from cacti to non-specialists, but the two are in completely different plant families separated by ~100 million years of evolution. Euphorbia species have milky toxic sap; cacti have clear sap.

Cultural and economic

Indigenous Mesoamerican cultures used Opuntia ([[prickly-pear|prickly pear]]) extensively: the pads as a vegetable (nopales), the fruit as a fruit (tunas), the cochineal-bug-infested pads as the source of a brilliant red dye (one of the most valuable Mesoamerican exports during the colonial period).

The peyote cactus ([[peyote|Lophophora williamsii]]) is the most-significant entheogenic cactus, central to Mexican Huichol religious practice and (under the Native American Church) to several US Indigenous traditions. San Pedro (Echinopsis pachanoi) is an Andean ceremonial cactus.

Cactus horticulture is a global industry. Mexico is the principal commercial producer of Opuntia and many ornamental cacti.

See also

Auto-generated from this entry’s typed relations: frontmatter, grouped by relation type so the editorial signal isn’t flattened.

  • Shares approach with: [[saguaro]]
  • Member of: [[plants]]

Sources

  • Wikipedia — Cactus

A plant entry in the 0mn1.one [[directory]].

What links here, and how

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Cultural

shares approach with

2 inbound links · 2 outbound